“It is devastating to hear of the passing of Joost,” said Howley.
“We all know his qualities on the pitch, he was a fantastic rugby player and for me was the best nine I played against. He had the ability to score tries from 5 or 70 yards, he was that good.
“I will never forget our game in Cardiff against South Africa in 1996. Joost scored three tries and was the different between the two sides.
“He was a world class nine who was respected throughout the rugby world.
“Away from the pitch we have enjoyed some very good times, reminiscing and catching up and that continued over the years, he even flew to Wales for my testimonial in 2005.
“I have been fortunate enough to play against him and enjoy his company off the pitch and it is tragic he has passed so young.
“Throughout his illness he remained the same character I always knew and he will be sorely missed.
“Our thoughts and best wishes are with all his family and friends at this time.”